diff --git a/src/build/targets.jam b/src/build/targets.jam
index 976422c82..876bab773 100644
--- a/src/build/targets.jam
+++ b/src/build/targets.jam
@@ -400,7 +400,7 @@ rule basic-target ( name : project
                         if dependency in [ feature.attributes $(p:G) ]
                         {
                             local g = [ generate-source $(p:G=) : $(properties) ] ;
-                            xproperties += $(p:G)$(g) ;
+                            xproperties += $(p:G)$(g) [ $(g).use-requirements ] ;
                         }
                         else
                         {
diff --git a/test/use_requirements.py b/test/use_requirements.py
index f398f6c63..b4b1193ad 100644
--- a/test/use_requirements.py
+++ b/test/use_requirements.py
@@ -21,6 +21,25 @@ t.run_build_system()
 
 t.run_build_system("--clean")
 
+# Test that use requirements on main target work, when they are referred using
+# 'dependency' features.
+t.write("project-root.jam", "import gcc ;")
+t.write("Jamfile", """
+    lib b : b.cpp : : : <define>FOO ;
+    exe a : a.cpp : <dependency>b ;
+""")
+t.write("b.cpp", "void foo() {}")
+t.write("a.cpp", """
+#ifdef FOO
+int main() {}
+#endif
+""")
+
+t.run_build_system()
+
+t.run_build_system("--clean")
+
+
 # Test that use requirement on project work
 t.write("Jamfile", "exe a : a.cpp lib/b ;")
 t.write("lib/Jamfile", """